Diversity and evolution of the ligule and contraligule in Scleria and related genera (Cyperaceae)
https://doi.org/10.1093/botlinnean/boaf109
Lucas Alves-dos-Santos, Bruno Edson-Chaves, Thomas Haevermans, Gladys F A Melo-de-Pinna
Botanical Journal of the Linnean Society, boaf109 (2025).
They tried to propose the new concept of ‘contraligule’ which has at least three conflicting definitions.