ペルー産カヤツリグサ科植物の再検討

A commented checklist and key for the genus Carex (Cyperaceae) in Peru

DOI: 10.3897/phytokeys.271.178418

Pedro Jiménez-Mejías, Pablo García-Moro, Andrea A. Camilo, Piero M. Mazzei,

Marcial Escudero, Santiago Martín-Bravo, Asunción Cano

PhytoKeys 271: 93 –144 (2026). 

56 species

six strictendemics

three subendemics

four first in Peru

Carex acutata 

Carex amicta 

Carex beckii 

Carex boliviensis 

Carex bonariensis: Absent 

Carex bonplandii 

Carex brachycalama 

Carex brehmeri 

Carex brevicaulis: Absent 

Carex camptoglochin 

Carex canoi: Endemic

Carex catamarcensis 

Carex chordalis 

Carex collumanthus 

Carex colombiana 

Carex crinalis 

Carex david-smithii 

Carex ecuadorica 

Carex enneastachya 

Carex erinacea: Absent 

Carex fecunda 

Carex gayana 

Carex goetghebeurii: ?

Carex guaguarum 

Carex haematopus 

Carex hamata 

Carex hebetata: Endemic

Carex herba-alpacae 

Carex hookeri: Absent 

Carex huancabambica 

Carex humahuacaensis 

Carex hypsipedos: Endemic 

Carex hypsipedos

Carex jamesonii 

Carex koyamae 

Carex laegaardii 

Carex lapazensis 

Carex lemanniana: ? 

Carex lepida 

Carex macloviana: Absent

Carex mandoniana 

Carex maritima: Absent

Carex melanocystis 

Carex melliza 

Carex meridensis 

Carex microglochin 

Carex ownbeyi 

Carex pachacutecii: ? 

Carex pachamamae: ?

Carex paramorum 

Carex perprava 

Carex peruviana: Absent 

Carex peucophila: Absent

Carex phalaroides: Absent

Carex phleoides: Absent 

Carex phylloscirpoides: ? 

Carex pichinchensis 

Carex polystachya: Absent 

Carex polysticha 

Carex porrecta 

Carex praegracilis: Absent 

Carex punicola 

Carex pygmaea 

Carex regina: subendemic

Carex roalsoniana 

Carex ros-desertum: Endemic

Carex ruthsatzae: ? 

Carex setifolia 

Carex sodiroi 

Carex tamana 

Carex tristicha 

Carex vallis-pulchrae: Absent

Carex viamontana 

ペルー -Google map-

https://maps.app.goo.gl/RUEPy3efoT5P8Cj87

韓国 光州市（全羅南道）のカヤツリグサ科植物

 Flora of Gwangju and the Jeollanam-do region

https://doi.org/10.11110/kjpt.2025.55.4.249

Eun-Mi SUN, Jin-Seok KIM, Hyoung-Tak IM

Korean J. Pl. Taxon 2025; 55(4): 249-274.

182,591 specimens:
collected from Gwangju and Jeonnam province

2,289 分類群
165 科 
770 属 
1,998 種 
50 亜種 
197 変種 
27 品種
17 雑種

<カヤツリグサ科 Cyperaceaee>
Carex fusanensis 부산사초 アカヒナスゲ　シノニム
Carex okamotoi 지리대사초　チイサンタガネソウ
Carex subebracteata var. leiosperma 지리실청사초　チイサンシバスゲ
Fimbristylis driziae 물하늘지기
Schoenoplectus nipponicus 물고랭이　シズイ

韓国 光州市（全羅南道） -Google map-
https://maps.app.goo.gl/CZkJrytzgzm3N4ku7

マダガスカルの国立公園のカヤツリグサ科植物

Patterns of Composition, Richness and Endemicity of Cyperaceae Across Open and Closed Habitats in Madagascar's Central Highlands

https://doi.org/10.1111/btp.70160

Fitiavana Rasaminirina, Jakub D. Wieczorkowski, Vonjison Rakotoarimanana, Valisoa Louisicaël Rafaralahy, Nivohenintsoa Rakotonirina, Hélène Ralimanana, Isabel Larridon

Biotropica, 58:e70160 (2026).

結果：
12属108種（固有種28種）
--乾燥地 (72種：個通種20種）
--湿地（60種：固有種7種）

調査地：マダガスカル
イザロ国立公園とイトゥルモ保護地域
標高 800m～2,700m alt.

地質サンプル：
20地点 (うち湿地6地点、乾燥地9点）

解析内容 (地質）:
pH, nitrogen, soil organic carbon, potassium and phosphorus

種リスト：
Abildgaardia ovata
Bulbostylis densa
Bulbostylis firingalavensis
Bulbostylis hispidula
Bulbostylis itremoensis
Bulbostylis micranthera
Bulbostylis perrieri
Bulbostylis schoenoides
Bulbostylis thouarsii
Bulbostylis trichobasis
Bulbostylis viguieri
Bulbostylis xerophila
Carex bathiei
Carex heterodoxa
Carex hirtigluma
Carex proxima
Carex pyramidalis
Carex renschiana
Coleochloa setifera
Costularia itremoensis
Costularia microcarpa
Cyperus albescens
Cyperus alleizettei
Cyperus alternifolius
Cyperus amabilis
Cyperus articulatus
Cyperus assimilis
Cyperus aster
Cyperus balfourii
Cyperus betafensis
Cyperus blastophorus
Cyperus brasiliensis
Cyperus brevifolius
Cyperus buchholzii
Cyperus chermezonii
Cyperus compressiformis
Cyperus cuspidatus
Cyperus cyperoides
Cyperus debilissimus
Cyperus denudatus
Cyperus dubius
Cyperus erectus
Cyperus esculentus
Cyperus flavescens
Cyperus haspan　コアゼガヤツリ　広義
Cyperus heterocladus
Cyperus hortensis
Cyperus hystricoides
Cyperus impubes
Cyperus intactus
Cyperus iria　コゴメガヤツリ　
Cyperus latifolius
Cyperus lipomexicanus
Cyperus longifolius
Cyperus longivaginans
Cyperus melanospermus
Cyperus mundii
Cyperus nigricans
Cyperus obtusiflorus
Cyperus papyrus　カミガヤツリ
Cyperus persquarrosus
Cyperus polystachyos
Cyperus procerus
Cyperus prolifer
Cyperus psammophilus
Cyperus pseudokyllingioides
Cyperus pseudovestitus
Cyperus pulchellus
Cyperus rhizomatosus
Cyperus rotundus　ハマスゲ
Cyperus sesquiflorus　タイトウクグ基本種
Cyperus unioloides
Cyperus varicus
Cyperus xerophilus
Eleocharis acutangula ミスミイ
Eleocharis dulcis　イヌクログワイ　
Eleocharis nigriscens
Eleocharis spiralis
Eleocharis variegata
Fimbristylis dichotoma　テンツキ　広義
Fimbristylis littoralis　ヒデリコ
Fimbristylis quinquangularis　イヌヒデリコ
Fimbristylis scabrida
Fuirena pubescens
Fuirena quercina
Fuirena stricta
Fuirena umbellata　ヒロハノクロタマガヤツリ　
Rhynchospora angolensis
Rhynchospora candida
Rhynchospora chinensis　イヌノハナヒゲ　シノニム
Rhynchospora corymbosa　ヤエヤマアブラスゲ　
Rhynchospora gracillima
Rhynchospora hildebrandtii
Rhynchospora holoschoenoides
Rhynchospora perrieri
Rhynchospora rubra
Rhynchospora rugosa　イヌノハナヒゲ基本種
Schoenoplectiella corymbosa
Schoenoplectiella erecta
Schoenoplectiella juncoides　イヌホタルイ　
Scleria angusta
Scleria boivinii
Scleria bulbifera
Scleria distans
Scleria greigiifolia
Scleria hilsenbergii
Scleria melanomphala
Scleria rutenbergiana

マダガスカル　イザロ国立公園 -Google map-
https://maps.app.goo.gl/iQCUK85mVPHFiXgf9

スゲにつく新種の病原菌

 Phylogenetic analyses, morphological characteristics, and host specificity reveal Anthracoidea koopmanii sp. nov. (Anthracoideaceae, Ustilaginales) on Carex oreophila (Cyperaceae) from Armenia

DOI: 10.11646/phytotaxa.740.3.1

TEODOR T. DENCHEV, CVETOMIR M. DENCHEV, 
HELENA WIĘCŁAW, VOJTĚCH MIŠEK, MARTIN KEMLER

Phytotaxa 740(3)(2026).

新種:
Anthracoidea koopmanii

Anthracoidea属の特徴:
スゲに寄生する特性を持つ

Carex alpestrisとC. nigraのレクトタイプ指定

Lectotypification of the names Carex alpestris All. and C. nigra Bellardi (Cyperaceae)

https://doi.org/10.3989/ajbm.662

Anales del Jardín Botánico de Madrid　8(2)(2025).

Alberto Selvaggi & Pedro Jimenez Mejias

韓国シロガヤツリ→コシロガヤツリ

Taxonomic reassessment of Korean specimens previously identified as Cyperus pacificus (Cyperaceae): Reporting C. michelianus from Korea

https://doi.org/10.11110/kjpt.2026.56.1.14
https://www.e-kjpt.org/journal/view.php?number=5155

Tae-Hun KIM , Eui-Kwon JUNG , Jung-Hoon LEE, Homerverge G. ONG , Chien-Ti CHAO , Young-Dong KIM

Korean Journal of Plant Taxonomy 2026;56(1): 14-24.

コシロガヤツリ Cyperus michelianus
韓国: 江原道以外の全域

Cyperus michelianus
Na-seon-huin-bang-dong-sa-ni 
나선흰방동사니

Cyperus pacificus var. pacificus
Huin-bang-dong-sa-ni
흰방동사니

Cyperus pacificus var. margoinflatus S. Fujii
Nal-gae-huin-bang-dong-sa-ni 
날개흰방동사니

ボリビアとブラジルからミカヅキグサ属２新種

Rhynchospora chiquicerradensis and Rhynchospora nucistipitata, new species from the Brazilian Cerrado and the Bolivian Chiquitano region

DOI: 10.11646/phytotaxa.739.2.6

PEDRO JOEL SILVA DA SILVA FILHO, SABRINA NICOLETTI, WILLIAM WAYT THOMAS

Phytotaxa 739(2) (2026).

Rhynchospora sect. Tenues

Rhynchospora chiquicerradensis
 from: the Chiquitano ecoregion of eastern Bolivia
 ttps://maps.app.goo.gl/Vfg8bkKVsjD9Ay3h9

Rhynchospora nucistipitata　
 from: the Cerrado domain of Brazil
 https://maps.app.goo.gl/sp6U2ff3GUGAdkHu9 

アリ散布型スゲ40種の散布様式と稔性の関係

Associations of dispersal traits with fecundity and clonal expansion among Japanese ant-dispersed sedges 

https://doi.org/10.1093/jeb/voag006

Koki Tanaka

Journal of Evolutionary Biology, voag006 (2026).

アリ散布型：
　長距離移動、大型エライオソーム、低稔性

重力・風散布型：
　茎が倒れて移動（近距離）、高稔性

クローン型：
　栄養繁殖（近距離）＞＞＞種子散布

ウクライナ ヤルネット保護地のカヤツリグサ科植物

Flora of the Yelanetsky Steppe Nature Reserve, Mykolaiv Oblast, Ukraine: a diversity and
structural analysis

DOI: 10.2478/environ-2025-0016

Galyna V. Drabyniuk, Oleksandr I. Shynder, Vitalii P. Kolomiychuk, Valeriia O. Konaikova

Environ. Socio.-econ. Stud. 13(3): 37-53 (2025).

Bolboschoenus glaucus
Bolboschoenus planiculmis　イセウキヤガラ
Carex hirta
Carex melanostachya
Carex michelii
Carex praecox　ヒメアサマスゲ　シノニム
Carex riparia
Carex spicata
Carex stenophylla
Carex supina　ホソバスゲ
Eleocharis palustris　クロヌマハリイ
Schoenoplectus tabernaemontani　フトイ
Scirpoides holoschoenus

ウクライナ　ムィコラーイウ州　ヤルネット保護地-Google map-
https://maps.app.goo.gl/ESpvEb95fzHD6wCm9

コウボウムギの米国での分布拡大報告

Biological Flora of Sand Dunes on the Atlantic Coastline of the United States: Carex kobomugi Ohwi, An Invasive Sedge

https://doi.org/10.2112/JCOASTRES-D-25A-00006.1

Richard Stalter, Robert I. Lonard

Journal of Coastal Research, 42(1): 122–126(2026).

日本→米国（帰化植物）
発見：1920年代　ニュージャージー州
現在：ノースキャロライナ州からマサチューセッツ州（米国右上）
拡大スピード：2 m/y (1平方メートルあたり140個体以上)

米国ノースキャロライナ州　-Google map-
https://maps.app.goo.gl/A15g5kCnZYFLzabg8

100年ぶりに発見されたハタガヤ属の一種

The Long-Lost Bulbostylis tenella (Cyperaceae): Rediscovery and Taxonomic Update with Lectotypification.

https://phytotaxa.mapress.com/pt/article/view/phytotaxa.737.5.2

Maciel-Silva, Juliene de Fátima, Larridon, Isabel, Prata, Ana Paula Do Nascimento,
Gil, André Dos Santos Bragança

Phytotaxa 737:(5): 276-282 (2026).

タイプ標本以降、117年ぶりに再発見
記載、レクトタイプ指定、絶滅危惧種指定、分布地図

ブラジル産ハリイ属２種の新絶滅危惧種指定

Eleocharis paranapiacabensis and E. raquellyrae: two new and endangered aquatic species of Eleocharis (Cyperaceae) from Brazil

DOI: 10.11646/phytotaxa.737.3.3

DIEGO C. FARIAS & MARCCUS ALVES

Phytotaxa 737(3) (2026).

Eleocharis ser. Maculosae (Cyperaceae):
  Eleocharis paranapiacabensis (Critically Endangered：CR(Critically Endangered)=絶滅危惧IA類)
  Eleocharis raquellyrae (Endangered：EN(Endangered)=絶滅危惧IB 類)

IUCN・環境省レッドリストカテゴリー説明
https://hirakawazoo.jp/wp/wp-content/themes/hirakawa/images/redlist.pdf

ブラジル -Google map-
https://maps.app.goo.gl/ZxoVqjRRPHWcDxhr5

カヤツリグサ属１類の生育地の違いによる分化

Molecular phylogeny reveals new morphological and ecological insights into the Cyperus margaritaceus–niveus complex

https://doi.org/10.1007/s00606-025-01983-w

Martin Xanthos, George Burton, Lisa Pokorny, Alice Merrall & Isabel Larridon 

Plant Syst Evol 312, 4 (2026). 

Cyperus margaritaceus–niveus complex (Cyperaceae):
  C4植物 (C. karlschumanii, C. kibweanus, C. ledermannii, C. margaritaceus, C. nduru, C. niveus var. leucocephalus, C. somaliensis, C. sphaerocephalus, and C. tisserantii)
  
分布：サブサハラ・アフリカ、マダガスカル

生育環境の違い:森林内と草地

系統解析に用いたDNA:核DNA
　本類が種間で類似しているため、市販のキット（Angiosperms353）を用いて353遺伝子から334遺伝子を用いて比較

ミヤマカンスゲ類の再検討と新提唱

Phylogeny and Taxonomy of Carex (Cyperaceae) in Japan II.
New Subsect. Multifoliae Separated from Subsect.
Pisiformes s.l. (Sect. Mitratae)

doi: https://doi.org/10.51033/jjapbot.ID0338

Jiro ODa, Shizuka Fuse, Jun YaMashiTa, Teruo KaTsuYaMa
and Minoru N. TaMura

J. Jap. Bot. 100(6): 489–514 (2025).
（Continued from Acta Phytotax. Geobot. 70(2): 69–85, 2019)

<方法>
DNA解析
・葉緑体DNA　plastid regions
 : atpBrbcL, rps3-rp116-rpl16, rps16 intron,  trnLtrnL-trnF
・核DNA　nuclear regions
 : ITS

形態
　痩果表面微細構造　Achene
　葉の断面　Leaf

<結果>
新亜節、新列、種の再定義を提唱

・亜節
1. C. subsect. Pisiformes (s.s.) sect. Mitratae
2. C. subsect. Multifoliae J.Oda & M.N.Tamura, subsect. nov.

・列
1. Carex ser. Multifoliae (J.Oda & M.N.Tamura) J.Oda & M.N.Tamura, stat. nov.
2. Carex ser. Imbecilles J.Oda & M.N.Tamura, ser. nov.
3. Carex ser. Dolichostachyae J.Oda & M.N.Tamura, ser. nov.

・列構成種
1. C. ser. Multifoliae:
 C. matsumurae
 C. toyoshimae
 C. augustini (C. toyoshimae var. augustini)
 C. atroviridis
 C. hachijoensis
 C. kuroshimensis

2. C. ser. Imbecilles:
 C. imbecillis
 C. pallidisquama

3. C. ser. Dolichostachyae:
 C. dolichostachya

・種再定義
1. クロシマナガボスゲ
Carex kuroshimensis J.Oda & M.N.Tamura (sp. nov.)
 Holotype: JAPAN. Kagoshima Pref.:
Kuroshima Isl., Mishima-mura, ca. 500 m alt.,
29 Apr. 2011, K.Yamawaki 16133 (KYO).
*Known only from  here
 Japanese name: Kuroshima-nagabo-suge (Hatusima 1991).

2. ウミノサチスゲ
Carex toyoshimae Tuyama
var. augustini (Tuyama) J.Oda, Katsuy. & M.N.Tamura, comb. & stat. nov.
Japanese name: Uminosachi-suge (Tuyama 1938).

3. アオミヤマカンスゲ
C. pallidisquama (Ohwi) J.Oda & M.N.Tamura (stat. nov.)
 Type: JAPAN. Mt. Katsuragi, Prov. Yamato (Nara Pref.), 4 May 1927, Z.Tasiro s.n. (KYO).
 Japanese name: Ao-miyama-kansuge (Akiyama 1955).

4. ナゴスゲ
C. dolichostachya var. connatisquama (T.Koyama) J.Oda &
M.N.Tamura (stat. nov.)
 (C. dolichostachya from Tokunoshima Isl., Kagoshima Pref.)
 Holotype: JAPAN. Okinawa Pref.: Mt. Nagodake, Okinawa Isl., S.Nakamine 988 (RYU, non vidi).
 Distribution: Endemic to Japan.
   Kagoshima Pref.: Okinoerabu Isl. and Tokunoshima Isl.
   Okinawa Pref.: Okinawa Isl. and Ishigaki Isl.
 Japanese name: Nago-suge (Matsumura 1905).