カヤツリグサ属１類の生育地の違いによる分化

Molecular phylogeny reveals new morphological and ecological insights into the Cyperus margaritaceus–niveus complex

https://doi.org/10.1007/s00606-025-01983-w

Martin Xanthos, George Burton, Lisa Pokorny, Alice Merrall & Isabel Larridon 

Plant Syst Evol 312, 4 (2026). 

Cyperus margaritaceus–niveus complex (Cyperaceae):
  C4植物 (C. karlschumanii, C. kibweanus, C. ledermannii, C. margaritaceus, C. nduru, C. niveus var. leucocephalus, C. somaliensis, C. sphaerocephalus, and C. tisserantii)
  
分布：サブサハラ・アフリカ、マダガスカル

生育環境の違い:森林内と草地

系統解析に用いたDNA:核DNA
　本類が種間で類似しているため、市販のキット（Angiosperms353）を用いて353遺伝子から334遺伝子を用いて比較

ミヤマカンスゲ類の再検討と新提唱

Phylogeny and Taxonomy of Carex (Cyperaceae) in Japan II.
New Subsect. Multifoliae Separated from Subsect.
Pisiformes s.l. (Sect. Mitratae)

doi: https://doi.org/10.51033/jjapbot.ID0338

Jiro ODa, Shizuka Fuse, Jun YaMashiTa, Teruo KaTsuYaMa
and Minoru N. TaMura

J. Jap. Bot. 100(6): 489–514 (2025).
（Continued from Acta Phytotax. Geobot. 70(2): 69–85, 2019)

<方法>
DNA解析
・葉緑体DNA　plastid regions
 : atpBrbcL, rps3-rp116-rpl16, rps16 intron,  trnLtrnL-trnF
・核DNA　nuclear regions
 : ITS

形態
　痩果表面微細構造　Achene
　葉の断面　Leaf

<結果>
新亜節、新列、種の再定義を提唱

・亜節
1. C. subsect. Pisiformes (s.s.) sect. Mitratae
2. C. subsect. Multifoliae J.Oda & M.N.Tamura, subsect. nov.

・列
1. Carex ser. Multifoliae (J.Oda & M.N.Tamura) J.Oda & M.N.Tamura, stat. nov.
2. Carex ser. Imbecilles J.Oda & M.N.Tamura, ser. nov.
3. Carex ser. Dolichostachyae J.Oda & M.N.Tamura, ser. nov.

・列構成種
1. C. ser. Multifoliae:
 C. matsumurae
 C. toyoshimae
 C. augustini (C. toyoshimae var. augustini)
 C. atroviridis
 C. hachijoensis
 C. kuroshimensis

2. C. ser. Imbecilles:
 C. imbecillis
 C. pallidisquama

3. C. ser. Dolichostachyae:
 C. dolichostachya

・種再定義
1. クロシマナガボスゲ
Carex kuroshimensis J.Oda & M.N.Tamura (sp. nov.)
 Holotype: JAPAN. Kagoshima Pref.:
Kuroshima Isl., Mishima-mura, ca. 500 m alt.,
29 Apr. 2011, K.Yamawaki 16133 (KYO).
*Known only from  here
 Japanese name: Kuroshima-nagabo-suge (Hatusima 1991).

2. ウミノサチスゲ
Carex toyoshimae Tuyama
var. augustini (Tuyama) J.Oda, Katsuy. & M.N.Tamura, comb. & stat. nov.
Japanese name: Uminosachi-suge (Tuyama 1938).

3. アオミヤマカンスゲ
C. pallidisquama (Ohwi) J.Oda & M.N.Tamura (stat. nov.)
 Type: JAPAN. Mt. Katsuragi, Prov. Yamato (Nara Pref.), 4 May 1927, Z.Tasiro s.n. (KYO).
 Japanese name: Ao-miyama-kansuge (Akiyama 1955).

4. ナゴスゲ
C. dolichostachya var. connatisquama (T.Koyama) J.Oda &
M.N.Tamura (stat. nov.)
 (C. dolichostachya from Tokunoshima Isl., Kagoshima Pref.)
 Holotype: JAPAN. Okinawa Pref.: Mt. Nagodake, Okinawa Isl., S.Nakamine 988 (RYU, non vidi).
 Distribution: Endemic to Japan.
   Kagoshima Pref.: Okinoerabu Isl. and Tokunoshima Isl.
   Okinawa Pref.: Okinawa Isl. and Ishigaki Isl.
 Japanese name: Nago-suge (Matsumura 1905).

中国産ヌカスゲ節３新種

Notes on Carex (Cyperaceae) from China (IX): three new species of section Mitratae s.l.

https://phytokeys.pensoft.net/article/101410/

Yi-Fei Lu, Zhao-Cen Lu, Yu-Hao Duan, Kun Zhang, Xiao-Feng Jin

PhytoKeys 225: 153-164 (2023).

1. Carex fatsuaniana X.F.Jin, Y.F.Lu & Z.C.Lu, sp. nov.
*urn:lsid:ipni.org:names:77318323-1
*similar: sect. Lageniformes

2. Carex damingshanica Z.C.Lu & X.F.Jin, sp. nov.
*urn:lsid:ipni.org:names:77318324-1
*similar: Carex breviscapa, C. rhynchachaenium

3. Carex radicalispicula Tang & F.T.Wang ex Y.F.Lu & X.F.Jin, sp. nov.
*urn:lsid:ipni.org:names:77318325-1
*With rhombic-ovoid nutlets abruptly contracted into a discoid-annulate style-base at the apex,

Cyperaceae Database

 Cyperaceae Database. 

https://cyperaceae.org/aphia.php?p=taxdetails&id=1847887

Carex daisenensis var. imbecillis (Ohwi) X.F.Jin & Y.F.Lu. 

Accessed at: https://cyperaceae.org/aphia.php?p=taxdetails&id=1847887 

on 2026-01-09

＊ヤワラミヤマカンスゲがダイセンスゲの変種

Jin, X. F., Lu, Y. F. (2024). 

Taxonomy of Carex sect. Mitratae s.l. (Cyperaceae).

 (Publication)

Carex daisenensis　ダイセンスゲ

　https://hoshinolab.main.jp/colorzukan/sugezk/tagyo/daisen/daisenp.htm

Carex multifolia var. imbecillis Ohwi　ヤワラミヤマカンスゲ

　https://umdb.um.u-tokyo.ac.jp/DShokubu/Carex_Japan/list/browserecord.php?-action=browse&No=TI00061659

高緯度湿地を生育地のヒトモトススキの遺伝的多様性

A Recent Lineage Split in Cladium mariscus Preceding the
Disjunction of Two Biodiversity Hotspots: Evidence From
Plastid Genomes

https://doi.org/10.1002/ece3.72456

Xin-Hui Qian, Liang-Liang Yue

Ecology and Evolution; 15:e72456 (2025).

葉緑体遺伝子の比較による第四期の気候変動の影響、
the role of Quaternary climate changes in driving divergence

葉緑体全遺伝子 (171,319 bp):
the complete chloroplast genomes (171,319 bp):
 IR region (IRA & IRB) (29,702bp)
 expanded LSC region
 exhibit strong A/T bias

採集地
from (high-altitude wetlands):
 雅魯蔵布江　the Yaluzangbu Valley (Bomi)
　　https://maps.app.goo.gl/1F6vCqrDGaTRxTih8
 横断山脈　　the Hengduan Mountains (Ninglang)
　　https://maps.app.goo.gl/tREBfoaaphKfA6he9

カザフスタン Kulaly島のカヤツリグサ科植物

Vegetation cover of the Kulaly Islands in the Kazakhstan region of the Caspian Sea

https://doi.org/10.31489/2025FEB4/76-91

M.S. agyndykova, G.G. Gasanova, A.A. Imanbayeva, A.E. Orazov

Fundamental And Experimental Biology, 30, 4(120): 76–91 (2025).

Carex physodes 
Bolboschoenus maritimus コウキヤガラ シノニム

the Kulaly Islands in the Kazakhstan -google map-
https://maps.app.goo.gl/rhMSdvcax5cHbLcP9

スゲ属の再定義

Nomenclature and typification of sectional names in Carex (Cyperaceae)

https://doi.org/10.1002/tax.70033

Pedro Jiménez-Mejías, James L. Reveal, Santiago Martín-Bravo, Juan Carlos Zamora,
Isabel Larridon, Anton A. Reznicek & Robert F.C. Naczi

Taxon (2025).

593 分類群
 33 属
  2 亜属
243/407 節 (illegitimate: 47) 
 32 亜節
  2 列
117 未ランク
250 年分の資料 (1753–2025)
100 学者
 54 タイプ標本の再定義
  4 新名
    Carex sect. Bracteosellae (Type: Carex bracteosa)
    Carex sect. Longirostrellae (Type: Carex longirostris (= Carex sprengelii))
    Carex sect. Stylosellae (Type: Carex stylosa)
    Carex sect. Triquetrellae(Type: Carex triquetra)

<Pick up> for Japanese

I. Carex subg. Siderostictae　タガネソウ亜属
I.A. “Carex sect. Siderostictae“　タガネソウ節
　Roalson et al. 2021　系統樹の番号
　https://doi.org/10.1111/jse.12722
  Circumscription: 27 species
Carex sect. Platyphyllae (Type: Carex siderosticta)
Carex sect. Hemiscaposae (Type: Carex scaposa)
Carex sect. Surculosae(Type: Carex surculosa)

VI.K. Carex sect. Phacocystis　アゼスゲ節
Carex sect. Forficulae　タニガワスゲ節
Type: Carex forficula タニガワスゲ

VI.X.4. Carex sect. Rhomboidales　ヒエスゲ節
Carex sect. Rhomboidales
Type: Carex chinensis Retz.

VI.X.5. Mitrata Clade　ヌカスゲ クレード
 Carex sect. Mitratae　ヌカスゲ節
 Carex sect. Praecoces　シバスゲ節
 Carex sect. Lageniformes　タイワンスゲ節

VI.X.6. Conica Clade ヒメカンスゲ クレード
 Circumscription: 48 species (Roalson et al. 2021)
Carex sect. Digamae 
Type: Carex digamma Nakai (= Carex conica Boott)